Cytosolic PLA 2 is required for CTL- Mediated immunopathology of celiac disease via NKG2D and IL-15

Fangming Tang, Zhangguo Chen, Cezary Ciszewski, Mala Setty, Jason Solus, Maria Tretiakova, Ellen Ebert, Jin Han, Anning Lin, Stefano Guandalini, Veronika Groh, Thomas Spies, Peter Green, Bana Jabri

Research output: Contribution to journalArticlepeer-review

79 Scopus citations

Abstract

IL-15 and NKG2D promote autoimmunity and celiac disease by arming cytotoxic T lymphocytes (CTLs) to cause tissue destruction. However, the downstream signaling events underlying these functional properties remain unclear. Here, we identify cytosolic phospholipase A 2 (cPLA 2) as a central molecule in NKG2D-mediated cytolysis in CTLs. Furthermore, we report that NKG2D induces, upon recognition of MIC + target cells, the release of arachi-donic acid (AA) by CTLs to promote tissue inflammation in association with target killing. Interestingly, IL-15, which licenses NKG2D-mediated lymphokine killer activity in CTLs, cooperates with NKG2D to induce cPLA 2 activation and AA release. Finally, cPLA 2 activation in intraepithelial CTLs of celiac patients provides an in vivo pathophysiological dimension to cPLA 2 activation in CTLs. These results reveal an unrecognized link between NKG2D and tissue inflammation, which may underlie the emerging role of NKG2D in various immuno- pathological conditions and define new therapeutic targets.

Original languageEnglish
Pages (from-to)707-719
Number of pages13
JournalJournal of Experimental Medicine
Volume206
Issue number3
DOIs
StatePublished - 16 Mar 2009
Externally publishedYes

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